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PHYLUM ARTHROPODA
PHYLUM ARTHROPODA
The phylum Arthropoda is the largest and most varied in the animal
kingdom. It includes well over one million described species. This
represents approximately three-quarters of all known biological
organisms, living or extinct. Countless arthropods remain undescribed, and the actual number of living species
could be as high as ten million or more. Some of the more well-known
arthropods include insects, crustaceans, and spiders. Arthropods are found in virtually
every known marine, freshwater, and terrestrial ecosystem, and vary tremendously in their habitats, life
histories, and dietary preferences.
Characteristics of Arthropods
All arthropods possess a stiff exoskeleton composed primarily of chitin
. In some species, lipids, proteins, and calcium carbonate may also
contribute to the exoskeleton. The external skeleton offers organisms
protection as well as support for the body. Its walls provide anchors
for the attachment of muscles. The exoskeleton is incapable of growth,
and is molted repeatedly during the growth of
the animal. This process is called ecdysis. Molting allows for rapid
growth until the newly secreted exoskeleton hardens.
Arthropod bodies are divided into segments. However, a number of
segments are sometimes fused to form integrated body parts known as
tagmata. This process of fusion is called tagmosis. The head, thorax,
and abdomen are examples of tagmata. Arthropods also have appendages
with joints. In early,
primitive anthropods, each body segment was associated with a single
pair of appendages. However, in most species some
appendages have been modified to form other structures, such as
mouthparts, antennae, or reproductive organs.
Some arthropods have highly developed sense organs. Most species have paired compound eyes
, and many also have a number of simpler eyes called ocelli.
Arthropods have an open circulatory system that
consists of a tube that is the heart and an open hemocoel
, the coelom of the animal, in which blood pools. Arthropods also have
a complete gut with two openings, the mouth and the anus.
Gas exchange in the phylum occurs in various ways. Some species have
gills, while others employ tracheae, or book lungs. The tracheal
respiratory system consists of external openings called spiracles that
are linked to a system of branched tubules which allow respiratory gases
to reach internal tissues. Arthropods are characterized by a brain as
well as a nerve ring around the area of the pharynx, in the oral cavity.
A double nerve cord extends backwards
along the ventral surface of the body, and each body
segment is associated with its own ganglion, or mass of nerve cells. In
most arthropod species, the sexes are separate. Fertilization usually
occurs internally, and most species are egg laying. While some species
exhibit direct development, in which eggs hatch as miniature versions of
adults, other species pass through an immature larval stage and undergo
a dramatic metamorphosis before reaching adult form.
Major Groups of Arthropods
Arthropods are divided into four subphyla. These are the Chelicerata,
the Crustacea, the Uniramia, and the Trilobita.
Subphylum Chelicerata
Subphylum Chelicerata comprises a major group within Phylum Arthropoda, including such animals as the arachnids (e.g., spiders and scopions), the extinct eurypterids, and the extant horseshoe crabs that are considered to be living fossils. These arthropods and their ancestral and extinct forms were and are mainly predators. Chelicerates are now predominently terrestrial animals, with most marine chelicerates, including all of the eurypterids, now extinct.
Their
name comes from their chelicerae, pointed appendages that grasp
food, that differ from the chewing
mandibles of most arthropods. Being unable to ingest solid food,
most Chelicerata either drink blood or spit or otherwise inject
digestive enzymes into their prey, and feed on the fluidized result.
Like all arthropods, chelicerates' bodies and appendages are covered
with a tough cuticle primarily composed of chitin and proteins
that chemically harden. Because this exoskeleton cannot stretch,
chelicerates have to molt in order to grow. Thus, they have to
molt the old, and await hardening of the new, during which time
they have reduced mobility and are otherwise more defenseless.
Subphylum Crustacea
Members
of Subphylum Crustacea (the Crustaceans) comprise a large
group of arthropods. The group contains familiar popular marine
food animals such as lobsters, crabs, shrimp. While mainly
found in salt and freshwater environments, there are also
terrestrial Crustacea such as woodlice and isopods. Crustaceans
have three distinct body parts: head, thorax, and abdomen
(also called a pleon). In some, the head and thorax are fused
to form a cephalothorax. They have two pairs of antennae on
the head, compound eyes, three pairs of mouthparts and a telson.
Crustaceans often have a thick carapace on the top (dorsal)
side that makes fossilization more likely; crabs and lobsters,
for example, have a thicker exoskeleton containing calcium
carbonate that is more readily fossilized.
Subphylum Uniramia
Uniramia (uni - one, ramus - branch, i.e. single-branches) is a group within the arthropods. In the past this group included the Onychophora, which are now considered a separate category. The group is currently used in a narrower sense.
Uniramia is one of three subphyla in Arthropoda classification suggested by Sidnie Manton. This classification divided arthropods into a three-phyla polyphyletic group, with phylum Uniramia including the Hexapoda (insects), Myriapoda (centipedes and millipedes) and the Onychophora (velvetworms). The discovery of fossil lobopods, determined to be intermediate between onychophorans and arthropods led to the splintering of the Lobopoda
and Onychophora into separate groups. This redefined the Uniramia as
strictly "true" arthropods with exoskeletons and jointed appendages.
Uniramians have strictly uniramous appendages.
Subphylum Trilobita
The subphylum Trilobita includes only extinct species found in fossil
form. The trilobites were a primitive group of marine species that was
particularly abundant during the Cambrian (570 million years ago) and
Ordovician (505 million years ago) periods. The group became extinct at
the end of the Permian (286 million years ago). Trilobites had
flattened, oval-shaped bodies. Most were a few inches long, although one
species is known to have attained a length of 0.6 meters (2 feet).
By
Reshma P
VASCULAR BUNDLES
VASCULAR BUNDLES
Plants have two systems for the transportation of substances -
using two different types of transport tissue. Xylem transports water
and solutes from the roots to the leaves, while phloem transports food
from the leaves to the rest of the plant. Transpiration is the process
by which water evaporates from the leaves, which results in more water
being drawn up from the roots. Plants have adaptations to reduce
excessive water loss.
Xylem and phloem
Plants have two transport systems to move food, water and
minerals through their roots, stems and leaves. These systems use
continuous tubes called xylem and phloem, and together they are known as
vascular bundles.
Plant stem
![]() |
| Stem – the xylem and phloem are arranged in bundles near the edge of the stem to resist compression and bending forces. |
Plant root
![]() |
| Root - xylem and phloem in the centre of the root to withstand stretching forces. |
Xylem
Xylem vessels are involved in the movement of water through a plant - from its roots to its leaves via the stem.
During this process:
- Water is absorbed from the soil through root hair cells.
- Water moves by osmosis from root cell to root cell until it reaches the xylem.
- It is transported through the xylem vessels up the stem to the leaves.
- It evaporates from the leaves (transpiration).
The xylem tubes are made from dead xylem
cells which have the cell walls removed at the end of the cells, forming
tubes through which the water and dissolved mineral ions can flow. The
rest of the xylem cell has a thick, reinforced cell wall which provides
strength.
Phloem
Phloem vessels are involved in translocation. Dissolved sugars, produced during photosynthesis,
and other soluble food molecules are moved from the leaves to growing
tissues (eg the tips of the roots and shoots) and storage tissues (eg in
the roots).
In contrast to xylem, phloem consists of columns of living
cells. The cell walls of these cells do not completely break down, but
instead form small holes at the ends of the cell. The ends of the cell
are referred to as sieve plates. The connection of phloem cells effectively forms a tube which allows dissolved sugars to be transported.
Transpiration
Water on the surface of spongy and palisade cells (inside the leaf) evaporates and then diffuses out of the leaf. This is called transpiration.
![]() |
| leaf |
More water is drawn out of the xylem cells inside the leaf to replace
what has been lost. Water molecules have a tendency to stick together –
so as water leaves the xylem to enter the leaf, more water is pulled up
behind it. This produces a continuous flow of water and dissolved
minerals moving up the xylem tube from the roots, up the stem, and into
the leaves. This is known as the transpiration stream.
Movement of water through the roots
The movement of water up the xylem means more water must be
drawn in through the roots from the soil. To do this, water passes from
root cell to root cell by osmosis.
As water moves into the root hair cell down the concentration gradient, the solution inside the root hair cell becomes more dilute. This means
that there is now a concentration gradient between the root hair cell
and adjacent root cells, so water moves from the root hair cell and into the adjacent cells by osmosis.
This pattern continues until the water reaches the xylem vessel within the root - where it enters the xylem to replace the water which has been drawn up the stem.
If the guard cells are turgid, then they curve forming ‘sausage-shaped’ structures with a hole between them. This is the stoma.
However, if the guard cells are flaccid due to water loss, they shrivel up and come closer together, closing the stoma. This is turn reduces the water loss due to transpiration, and can prevent the plant from wilting.
By
Vinesh S
![]() |
| The pathway of water across a root |
This pattern continues until the water reaches the xylem vessel within the root - where it enters the xylem to replace the water which has been drawn up the stem.
Factors that affect transpiration rate
Light
Transpiration increases in bright light. The stomata open wider to allow more carbon dioxide into the leaf for photosynthesis. More water is therefore able to evaporate.
Temperature
Transpiration is faster in higher temperatures. Evaporation and diffusion are faster at higher temperatures.
Wind
Transpiration is faster in windy conditions. Water vapour is removed quickly by air movement, speeding up diffusion of more water vapour out of the leaf.
Humidity
Transpiration is slower in humid conditions. Diffusion of water vapour out of the leaf slows down if the leaf is already surrounded by moist air.
Factors that speed up transpiration will also increase the rate of water uptake from the soil. If the loss of water is faster than the rate at which it is being replaced by the roots, then plants can slow down the transpiration rate by closing some of their stomata. This is regulated by guard cells, which lie on either side of a stoma.
![]() |
| Plants can slow down the transpiration rate by closing some of their stomata |
If the guard cells are turgid, then they curve forming ‘sausage-shaped’ structures with a hole between them. This is the stoma.
However, if the guard cells are flaccid due to water loss, they shrivel up and come closer together, closing the stoma. This is turn reduces the water loss due to transpiration, and can prevent the plant from wilting.
By
Vinesh S
IMMUNOLOGY
IMMUNOLOGY
Anything that causes an immune response is called an antigen. An antigen may be harmless, such as grass pollen, or harmful, such as the flu virus. Disease-causing antigens are called pathogens. The immune system is designed to protect the body from pathogens.
In humans, the immune system begins to develop in the embryo. The
immune system starts with hematopoietic (from Greek, "blood-making")
stem cells. These stem cells differentiate into the major players in the
immune system (granulocytes, monocytes, and lymphocytes). These stems
cells also differentiate into cells in the blood that are not involved
in immune function, such as erythrocytes (red blood cells) and
megakaryocytes (for blood clotting). Stem cells continue to be produced
and differentiate throughout your lifetime.
Hematopoietic stem cells produce cells in blood and lymph
By the time a baby is born, the immune system is a sophisticated
collection of tissues that includes the blood, lymphatic system, thymus,
spleen, skin, and mucosa. The immune system is typically divided into two categories--innate and
adaptive--although these distinctions are not mutually exclusive.
Innate immunity
Innate immunity refers to nonspecific defense mechanisms that come into
play immediately or within hours of an antigen's appearance in the body.
These mechanisms include physical barriers such as skin, chemicals in
the blood, and immune system cells that attack foreign cells in the
body. The innate immune response is activated by chemical properties of
the antigen.
Adaptive immunity
Adaptive immunity refers to antigen-specific immune response. The
adaptive immune response is more complex than the innate. The antigen
first must be processed and recognized. Once an antigen has been
recognized, the adaptive immune system creates an army of immune cells
specifically designed to attack that antigen. Adaptive immunity also
includes a "memory" that makes future responses against a specific
antigen more efficient.
The cellular system
- T-cells differentiate in the thymus, and have a specific receptor for a fragment of antigen..
- Cytotoxic T-cells contain a surface protein called CD8 and destroy pathogen infected cells, cancer cells, and foreign cells (transplanted organs).
- Helper T-cells contain a surface protein called CD4 and regulate both the cellular and humoral immune systems. This regulation reduces autoimmunity.
- Autoimmune disease- self immunity. Some examples include rheumatic fever, rheumatoid arthritis, ulcerative colitis, myasthenia gravis, etc.
Immunological response
The graph shows a very important feature of the immune response. When first exposed to antigen "A",
we begin to make low levels of antibody in about a week However, a second exposure to antigen "A"
produces a much faster response,
and several orders of magnitude higher levels of antibody. The ability of antibody to
bind antigen also increases dramatically in the secondary response. Injecting a new antigen "B" with "A"
shows that a memory or prior exposure is required for the accelerated response. The memory of
antigen and the stimulated response is the basis for success in vaccination programs.
The Clonal Selection Theory
- The immune systems produces Billions of kinds of B-cells each making one kind of antibody receptor.
- The presence of antigen leads to the proliferation and differentiation of clones that have antibody capable of binding the antigen. In the diagram the "green" antigen binds to the green antibody on a B-cell. The color code means that only this antibody receptor on the cell binds free antigen.
- The "green" helper T-cell must give a stimulatory signal to allow a particular B-cell to be selected. This step allows a regulation or control of the process.
- The antigen driven selection produces memory cells and plasma cells secreting antibody capable of binding the original selecting antigen with high affinity..
- If antigen appears in the organism a second time, then the memory cells are already present at high levels, and produce a more rapid and much stronger immune response.
By
Aparna S M
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